Towards The Optimization Of Resistance Gene Analog Discovery In Solanum Species Quirin, E.A. and J.M Bradeen

Plant & Animal Genome XIII (San Diego, CA Jan 2005)

Resistance gene analogs (RGA) are potential gene fragments that contain structural motifs, such as the NBS and LRR regions, that are common to most cloned R genes. Recovery of RGAs using degenerate primer PCR was first described for cultivated potato, and has become a popular method for recovering RGAs from many dicot and monocot plant species. In this study, we are applying the degenerate primer PCR approach to the disease resistant, wild potato species, Solanum bulbocastanum. We aimed to compare total genomic DNA versus pooled BAC DNA as template for RGA recovery in S. bulbocastanum. From total genomic DNA, we observed an excess of PCR fragments originating from highly repetitive genomic regions. Normalization during BAC library construction enriches for euchromatic, gene-rich regions, potentially minimizing the highly repetitive sequences available for amplification. We compared total genomic DNA versus pooled BAC DNA RGA reactions for the recovery of R gene like fragments and for the overall amplification of repetitive elements using DNA hybridization. We further evaluated the pooled BAC strategy by estimating the frequency of fragments originating from BAC vector and E. coli DNA. Finally, we determined the minimum genomic equivalents needed to produce a consistent banding pattern by varying the number of BAC clones of each pool.